"During the demonstration, called Project Riot, an F-35 detected a long-range missile launch with its onboard sensors and shared the information through the U-2 to the air defence commander on the ground, enabling the commander to quickly make the decision to target the threat," the release said.
Lockheed characterised the test as a "ground-breaking" demonstration of "multi-domain operations and the secure distribution of sensitive information across multiple platforms."
This test showed the "next-level" of connectivity intended to reduce the data-to-decision timeline from minutes to seconds, according to the release.
The demonstration also leveraged the F-35 sensor data for use in missile defence, the release added.